From the BlackBerry's Home screen, go into the Messages application, then Click the Scroll Wheel or [Menu] button and select Options --> Email Reconciliation.
If you have more than one email account being delivered to the device, hit [Space] until Message Services is set to "Desktop", then highlight the field to the right of "Delete On:" until it says "Mailbox & Handheld". Hit [Esc] and save the changes.
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